Honored for their “cultural and natural heritage of outstanding value to humanity,” Spain boasts a whopping 45 UNESCO sites—the third most after Italy and China. While Barcelona, Seville, and Granada are noteworthy classics, consider trading some of the usual tourist stops for these lesser known destinations.

DOÑANA NATIONAL PARK: A NATURAL PARADISE

Nestled in the middle of three southern provinces, Doñana National Parkis one of Europe’s most significant conservation areas. The 209-square-mile biosphere reserve is a jigsaw puzzle of lagoons, marshes, lakes, cliffs, woodlands, and long stretches of pristine beaches untouched by human development.

With miles of trails throughout the park, visitors can spend hours hiking, biking, and birdwatching, with hundreds of species to admire.